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Showing posts with label apical dominance. Show all posts
Showing posts with label apical dominance. Show all posts

Thursday, January 31, 2013

Gardening Study School: Pruning Handout

On both these illustrations, you can clearly see
the Bark Branch Ridge and where to cut
in relationship to it.

General Notes:

Pruning involves the selective removal of specific plant parts for the benefit of the entire plant.
  • usually involves the removal of branches
  • can include removal of roots, seed pods, or flower buds.
  • many plants will grow and flower with little pruning provided they are given sufficient room to grow. (Don't prune it unless needed!)

    Pruning is
  • frequently misunderstood and incorrectly practiced.
  • Common to wait until a plant has out-grown its intended space before deciding to prune.
  • others incorrectly view pruning as an annual spring ritual that must be done --- sometimes when it is not needed.

    Pruning should be viewed
  • as a regular part of maintenance
  • not a remedial correction of neglected problems.
Pruning to maintain plant health includes the elimination of dead, dying, or diseased wood. A dying branch or stub can be an entry point for insects or diseases that could spread to other parts of the plant. When removing diseased wood it is important that the cut be made with a sterile blade into healthy wood. Dip the blades in a disinfectant between each cut or you could spread the disease. (choice of disinfectant – bleach? Listerine? Alcohol? )

Pruning special effect as is the case with espalier, or tree form plants.

Pruning is sometimes undertaken to help a plant recover from transplant shock or from construction damage.
Trees are pruned to eliminate low-growing branches that interfere with traffic or to thin the branches (lacing out) to allow more sunlight to reach plants that are growing under them.

Pruning can enhance the flowering display of some plants. As a plant matures it produces more but often smaller flowers. Pruning reduces the amount of wood and so diverts energy into the production of fewer but larger flowers.

Pruning-off faded flower clusters of crapemyrtle will stimulate the production of new flower buds and a second round of color.
Pruning can be used to produce longer, stronger stems, more colorful stems , or more attractive fruiting.

Response to pruning
A basic understanding of how a plant responds to pruning will help you do a better job. The terminal bud on a shoot produces a hormone (auxin) that inhibits the development of lateral buds along the shoot. When the terminal bud is removed the lateral buds near the cut (6 to 8 inches below) become active and grow. Regrowth on a branch with a 45 to 60 degrees angle will develop farther down the shoot.

When cutting back to a bud, make the cut at a slight angle just above the bud.

NOTE: The angle will allow moisture to flow off the cut. Avoid making the cut at a sharp angle because it will produce a larger wound. When you cut a branch back to the main trunk or to a lateral branch or bud the wound will heal quicker. When you prune leaving a stub the cut is slower to heal. The dead stub is an entrance point for disease and insects.

While pruning is done to reduce the overall size of a plant, it should be remembered that the growth of shoots near the pruning cut are invigorated. Strong shoots should be moderately pruned and weak shoots should be severely pruned. Severely pruning strong shoots will cause vigorous growth.

Correct pruning is the selective removal of branches while maintaining the natural shape of the plant. It is not the same as shearing in which the ends of most, if not all, branches are removed.


A good pruning job should not be conspicuous.
Before starting consider the natural form of the plant; choose branches that do not fit the natural form for removal. Each plant species has an individuality that distinguishes it from other plants.

Shearing increases maintenance; any new growth that occurs will seem out of place and extend beyond the artificial shape that has been created. Pruning to maintain the natural shape is less formal but looks better. New shoot growth tends to blend in and the time between pruning is extended.

Types of pruning

Five basic techniques are used for pruning shrubs:

pinching
heading back
thinning
pruning for renewal
and shearing.
Some plants require more of one method than another, but good pruning is usually a combination of several methods.

Pinching - is the removal of the terminal portion of a succulent, green shoot before it becomes woody and firm. Pinching can greatly reduce the need for more dramatic pruning later on. Whenever (except late summer) you see a shoot becoming excessively long simply pinch or cut the shoot to reduce its length and to promote side branching. Long, vigorous shoots should be cut back into the canopy instead of cut at the outer limits of the existing foliage.

Heading back - involves removing the terminal portion of a woody branch by cutting it back to a healthy bud or branch. Heading back will stimulate shoot growth below the cut thus making the plant more dense. The shape of the plant can be influenced by cutting to inward or outward growing buds. The top bud should be located on the side of the branch that faces the direction new growth is desired. Some plants will have two buds opposite each other on the stem. When such stems are cut, remove one of the buds if you need to control the direction of new growth. If both are allowed to grow, a forked and often weak stem may develop. Repeated heading back with no thinning cuts results in a top heavy plant. Dense top growth reduces sunlight and results in the loss of foliage inside the plant canopy.

Thinning - is the least conspicuous method of pruning and results in a more open plant without stimulating excessive new growth. Considerable growth can be cut without changing the plant’s natural appearance or growth habit. With thinning cuts a branch is cut off at its point of origin from the parent stem, to a lateral side branch, to the “Y” of a branch junction, or at ground level. A good rule-of-thumb is to prune to a lateral that is one-third the diameter of the branch being removed. Thin out the oldest and tallest stems first, allowing vigorous side branch development. This method of pruning is best done with pruning shears, loppers, or a saw --- not hedge shears.

Renewal pruning (rejuvenation) - involves removing the oldest branches of a shrub by pruning them near the ground, leaving only the younger, more vigorous branches which may also be cut back. Small stems (less than pencil size) should be removed.

A variation of renewal pruning involves cutting all branches back to a predetermined height each year. Butterfly bush is often pruned back to woody framework. With time the framework becomes congested and will require some slight thinning.

Shearing - involves cutting the terminal of most shoots with shearing or hedge clippers. This method should not be used on foundation plants but should be restricted to creating formal hedges. Shearing destroys the natural shape of the plant. It causes a thick profusion of growth on the exterior of the plant that excludes light form entering the center of the plant (Figure 15-30). Foliage on the interior of the plant dies. The natural renewal growth from within the plant in prevented.

Time of pruning
Light corrective pruning can be done any time of the year. Occasional, light pruning will reduce the need for severe pruning which might weaken the plant and reduce its natural beauty. When severe pruning becomes necessary it should be undertaken in late winter to early spring.

Woody plants heal pruning cuts by producing rolls of callus that gradually grow over the wound. Internally they compartmentalize or wall-off the damaged tissue from healthy wood. These responses to wounding occur more rapidly just prior to the onset of new growth in the spring and just after maximum leaf expansion in June. The time of pruning will also affect the amount of new growth produced. Plants respond to late winter and early spring pruning by producing vigorous growth to replace the removed wood.
A. When How and Why

  1. Pruning normally takes place in the winter months for deciduous trees
  2. Sap flow in trees is reduced and the garden calendar is much reduced
  3. The 'ideal' window in our world is from the ML King holiday to Valentines Day – it is getting earlier! For citrus, and other non-deciduous trees, this is not true – prune them after they have flowered or fruited regardless the time of year.
  4. Trees and some shrubs left to their own devices outgrow the space we've allotted for them, but also grow into a less manageable mass if proper pruning is not done.
    PLEASE NOTE: as you drive around and see government workers and crews pruning trees and shrubs, you can bet money that most of that work is incorrect! Pruning is an art and a science and both have been thrown out the window in a headlong race to hire the lowest bidder.
B. Tools
  1. Pruners (a bypass or secatuers not anvil style pruners!), loppers, saw(s), knife, pole saw and pole pruner
    a. pruners are your best weapon – cleanest cut
    b. saws are your second best cut
    c. lopper cuts often need to be cleaned up with a knife
    d. pole pruners and pole saws are difficult to control for correct cuts.
  2. Sharp and clean – disinfected too
    a. sharpening demo – hand held stone to the belt sharpener
    b. disinfectants – alcohol, 10% bleach solution, Listerine
  3. A ladder for some trees
    a. not your household ladder!
    b. tripod, orchard ladder
  4. Dress appropriate to the job at hand
    a. gloves
    b. not a lot of blowy clothes
C. The Principles of Pruning
  1. Cut any damaged, diseased or dead wood first.
  2. Cut out crossing wood
  3. Open the center of the plant to allow light and air flow.
  4. Prune to shape, if any more pruning is needed.
  5. Cut to a node, paying attention to the apical dominant points, through the bark branch ridge and make the cut as clean as possible.
  6. Stop pruning and step back frequently to assess the progress. 'Ask' the tree/shrub what shape it needs to be. DO NOT RUSH. Do not get frozen with anxiety, but don't blithely hack the plant apart.
  7. Depend on your pruners mostly with the pocket saw as your second most used tool – use the lopper only as necessary – and use the pole tools the least!
  1. Demonstration, indoors and outdoors. Student participation encouraged.


Saturday, January 23, 2010

Definitions and Terminology for Propagation

Buds on a named-variety of Ceanothus - all Ceanothus hybrids are propagated by cuttings; the hybrid can't be maintained any other way because the crosses aren't stable.

Apical dominance Apical dominance is phenomenon where the tip of the main trunk or branch bears the dominant meristem. The tip is apical meristem. This allows the convenience of the tip to grow fast and avoid being shadowed by branches. If the dominant meristem is removed, one of the nodes left on the branch or trunk will assume dominance, usually the topmost bud/node. 

Apical meristem The growing tip composed of completely undifferentiated meristematic tissue, these are the buds (nodes) and growing tips of roots in plants.

Asexual reproduction (also known as vegetative reproduction) Any reproduction in plants that does not involve meiosis or fertilization; there is only one parent contributing genetic material to the resulting offspring – they are, in effect, “cloned.”

Bud See node.

Callus
The mass of parenchyma cells that develops from and around wounded plant tissues in order to diminish evaporation from the wound and initiate healing. In grafting it occurs at the junction of a graft union, arriving from living cells of both scion and stock. The production and interlocking of these parenchyma (or callus) cells constitute one of the important steps in the healing process of a successful graft.

Cambium A thin meristematic tissue of the plant located between the bark and the wood. Its cells are are capable of dividing and forming new cells. For a successful graft union, it is essential that the cambium of the scion be placed in close contact with the cambium of the stock.

Clone A population of genetically identical cells or individuals. Such a population is obtained by mitotic division or by asexual reproduction.

Distal The distal end of either the root or the shoot is that furthest from the stem-root junction of the plant and nearest to the tip of the shoot or root. Alt. proximal. See polarity.

Dormant/dormancy (buds) A bud which have stopped its development for a period due to unfavorable environmental conditions (e.g. a dry or cold season). A dormant bud will sprout as a response to improved growth conditions or a “biological clock” e.g. longer days.

Grafting The connection of two pieces of living plant tissue in such a manner that they will unite and subsequently grow and develop as one plant.

Grafting wax Substance applied on the graft union in order to minimize desiccation and exclude water access.

Graft union The site of the grafted plant where the scion and the root stock are united.

Hedging Trimming trees and keeping them low in order to overcome or bypass the poor rooting and often poor form of cuttings from old trees. Cuttings from hedged plants tend to maintain their young physiological age.

Hormone A substance that has a marked effect on a specific plant part and produces this effect when present in very low concentrations, e.g. promotion of root, shoot or flower development. Hormones are produced within the plant but artificial synthesized plant hormones applied to the plant part have the same effect.

Incompatibility (graft incompatibility) Inability of the stock and the scion to form or maintain a union that will result in the desired plant growth.

Meristem (meristematic tissue) This is tissue in all plants consisting of undifferentiated cells (meristematic cells) and found in zones of the plant where growth can take place. Grafting and cuttings rely on this phenomenon of plant physiology to achieve results.

Node The horticultural term for a point of the stem from which one or more leaves arise or can arise. In the mature stem the nodes are usually well separated by internodes which elongate during growth. Apical meristem is the botanical term. Nodes may be dormant and only visible as a slight swelling along a stem.

Scion An aerial plant part, often a branchlet, that is grafted onto the root bearing part (stock/root-stock) of another plant.

Sexual reproduction  Plants reproduce sexually in a number of ways, but for the purpose of this course where we are dealing with primarily higher (flowering) plants, sexual reproduction takes place through pollen from a male flower being transferred to the female flower and the resulting fertilization resulting in the growth of seeds with the following generation having genetic traits of both parents.

Strike Slang for a cutting that has rooted. Occasionally used as a verb for the act of making a cutting.

Stock/root stock The lower portion of the graft, which develops into the root system of the grafted plant. It may be a seedling, a rooted cutting or a layered plant.

Take A slang term for the phenomenon of the successful union of stock and scion and growth of the grafted plant.

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